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Rufus recognised a more modern eye, with conjunctiva, extending as a fourth epithelial layer over the eye. Rufus was the first to recognise a two chambered eye - with one chamber from cornea to lens (filled with water), the other from lens to retina (filled with an egg-white-like substance). Galen remedied some mistakes including the curvature of the cornea and lens, the nature of the optic nerve, and the existence of a posterior chamber. Though this model was roughly a correct but simplistic modern model of the eye, it contained errors.
Yet it was not advanced upon again until after Vesalius. A ciliary body was then discovered and the sclera, retina, choroid and cornea were seen to meet at the same point. The two chambers were seen to hold the same fluid as well as the lens being attached to the choroid. Galen continued the notion of a central canal, though he dissected the optic nerve, and saw it was solid, He mistakenly counted seven optical muscles, one too many. He also knew of the tear ducts.
Even the pain that so often accompanies errors of refraction is never caused by the same strain that causes these errors. This layer is composed of minute rodlike and conical bodies which vary in form and are distributed very differently in its different parts. They are directed to look at such an object at the distance the hypermetropia is lessened, but pain and discomfort may be increased. Even total blindness lasting a few hours has been produced. If it does not appear equally black to you under all these conditions, therefore, you may know that your sight is imperfect. Many children could read one Snellen test card: with perfect sight, while unable to see a different.
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